Do Aeonium, Agave, Aloe, and Echeveria show watermarks like Sempervivum can show?

I have always been curious, which succulents can or cannot have watermarks and how impressive or 'dull' they can be.







Most sempervivum watermarks are wavy patterns, heuffelii can have dots watermarks, but interestingly enough they are unconvincing and rare and only on the outer leaves, in my case almost all heuffelii cultivars that watermark look like in this image, far from the impressive wavy patterns shown above with non-heuffelii sempervivum.

I'm not sure exactly what defines a watermark, but several glaucous agaves have something the agave people call crossbanding (among other names), which looks similar to the first couple of pictures.

Searching around I have found the technical explanation according to Kevin Vaughn with regards to sempervivum.

JungleShadows said: Yes the watermarks are from uneven distribution of surface waxes that causes the underlying anthocyanin colored cells beneath to have different shades. Think about adding candle wax over a basic clear color in different amounts and you have an idea as to how the colors can be affected. It adds a lot of distinction and beauty to those that exhibit these characteristics.

Kevin

I assumed that the uneven wax distribution would be cause by water and temperature (possibly light)
But now I am not sure if water is involved at all based on the explanation.

In my mind (mostly my imagination) the patterns on those agaves might have to do with irregular growth patterns. Any given leaf may stop and start (based on water availability or some kind of natural program) as it extends to reach full length. In the process the production of the waxy farina is somehow not well coordinated with growth, resulting in an accumulation during those pauses or upon stopping or restarting. But that is just a guess. I really don't know why it happens, or if there is anything we do as gardeners to promote this curious pattern. If water has anything to do with it, the mechanism would relate to how water availability governs the starts and stops. I am guessing the watermark term is just an analogy to something visually similar that may happen with paper and ink.